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Introduction to Evolution and Natural Selection
Evolution refers to the changes in the heritable characteristics of biological populations over successive generations. It is one of the most important frameworks in biology, explaining the incredible diversity of life on Earth, from microscopic bacteria to towering redwood trees.
Natural selection is a key mechanism driving evolution. It acts on the variations within populations, favoring genetic traits that confer survival and reproductive advantages. Over many generations, these traits become more common, leading to evolutionary changes.
A solid grasp of evolution and natural selection is essential for the AP® Biology exam. Not only are these theories fundamental to modern biology, but they also appear frequently in exam questions and are often central to understanding genetics, ecology, and population dynamics.
Understanding Evolution and Natural Selection
Evolution, in simple terms, is the process by which different kinds of living organisms develop and diversify from earlier forms. This concept provides a unifying theme in biology, showing how species can adapt over time due to various factors, including mutations, genetic drift, and gene flow.
Natural selection is the differential survival and reproduction of individuals based on their inherited traits. Organisms with traits better suited to their environment tend to survive longer and produce more offspring—thus passing their advantageous genes onward.
Charles Darwin, in his seminal work “On the Origin of Species,” described how natural selection drives evolutionary change. His observations of finches in the Galápagos Islands helped illustrate how specific traits, such as beak size and shape, could be favored in different environmental conditions.
The Four Principles of Natural Selection
A. Variation
Populations exhibit genetic differences. These variations can result from random mutations, gene shuffling during sexual reproduction, and other sources of genetic diversity. Mutations can introduce new alleles, while recombination (during meiosis) reshuffles existing alleles into new combinations, fueling the raw material upon which natural selection acts.
B. Competition
Because food, space, and other resources are finite, organisms must compete to survive. Only some individuals will secure enough resources to thrive and reproduce. Environmental conditions, such as climate or the presence of predators, can dictate which traits are most advantageous in a population at any given time.
C. Differential Survival
Individuals that possess traits better suited to the environment—such as camouflage, disease resistance, or efficient metabolism—are more likely to survive. Consider the classic example of the peppered moth in industrial areas of England. As pollution darkened tree bark, darker moths escaped predation more easily, leading to a shift in the common coloring of the moth population.
D. Reproduction
“Fitness” in an evolutionary context is measured primarily by reproductive output—put simply, who leaves more offspring that survive to reproduce. Traits that help an organism survive (and thus reproduce) are passed to the next generation, gradually becoming more prevalent.
Natural Selection’s Effect on Populations
A. Evolutionary Fitness
Biologists often measure fitness as the relative number of viable, fertile offspring an individual produces compared to others in the same population. Fitness can be quantified with counts of offspring or by examining which genotypes increase in frequency over time. For example, antibiotic-resistant bacteria reproduce more successfully in an environment with antibiotics, indicating high fitness levels for the resistant genes.
B. Environmental Influence
- Biotic and Abiotic Factors Affecting Evolution
- Predator-prey relationships, diseases, resource availability, temperature, and climate change are examples of biotic (living) and abiotic (nonliving) factors that shape the direction of natural selection.
- Stability vs. Fluctuations in Environments
- Stable environments often favor well-adapted traits over the long term, while changing environments can drive rapid evolutionary shifts.
- How Genetic Variations Are Selected Differently Across Generations
- If an environment changes (for example, a shift in climate), different variations may become favorable, potentially altering allele frequencies in the population over multiple generations.
Real-World Applications and Examples
A. Case Studies of Natural Selection
- Antibiotic Resistance in Bacteria
- Bacterial populations exposed to antibiotics quickly evolve resistance as individuals with resistance genes survive and pass these genes on. This is a pressing global health example of natural selection in action.
- Darwin’s Finches
- These finches on the Galápagos Islands provided Darwin insight into how slight variations in beak shape and size offered survival advantages in different environments, driving speciation.

B. Illustrations to Visualize Concepts
Imagining or sketching population changes as environmental pressures shift can clarify how natural selection shapes species. Graphs showing how a trait frequency changes over generations can make the concept more concrete.
Practice Problems
- Suppose a hypothetical population of moths has two color variants: light and dark. If the environment becomes polluted and tree bark darkens, which variants will likely have higher fitness? Why?
- Use Hardy-Weinberg analysis to determine how selection pressure on one genotype impacts the overall allele frequencies in a population over time.
Conclusion
We’ve discussed how evolution is driven by natural selection—relying on genetic variation, competition for resources, differential survival, and reproductive success—to shape species over time. These concepts form the foundation of modern biology and will appear repeatedly in AP® Biology. Understanding them thoroughly will boost confidence and exam performance. Continue exploring by diving into more real-world examples, reviewing data on how populations shift, and practicing questions that apply these principles to various scenarios.
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